Skip to main navigation menu Skip to main content Skip to site footer

On the Rediscovery of Pellaea Calomelanos in Central Himalaya with a Note on Distribution


Among the rare ferns of India, Pellaea calomelanos (Sw.) Link has been re-discovered from Kumaon region of central Himalaya after a gap of nearly 100 years. In the intervening years this rare fern was not collected from any other parts of Himalaya or India. A detailed note on its distribution in India, and other parts of the world along with a note on colonisation is elaborated.


Kumaon, Distribution, Pellaea calomelanos, Re-Discovery.



Download data is not yet available.


  1. BARRINGTON, D.S. 1993. Ecological and historical factors in fern biogeography. Journal of Biogeography 20: 275-280.
  2. BEDDOME, R.H. (1863-1864). The Ferns of British India. Madras: Gantz Brothers (1863-1864). 1-38, t. 1-110 (1863); 3-88. t. I11-271 (1864). Reprint: Today & Tomorrow's Printers & Publishers. New Delhi (1970).
  3. BEDDOME, R.H. 1883. Handbook to the Ferns of British India: Ceylon and the Malay Peninsula; with Supplement (1892). Thacker, Spink, and Co., Calcutta, India., xvi/500/110/(4) pp., 300 ill.
  4. CHANDRA, S. 2000. The ferns of India (Enumeration, Synonyms & Distribution). International Book Distributors, Dehradun, India. 459pp.
  5. CLARKE, C.B. 1880. A Review of the Ferns of Northern India. Trans. Linn. Society Series 2: Botany, Vol. 1. London. (Reprinted in Delhi 1973).
  6. DHIR, K.K. 1980. Ferns of North Western Himalayas. Bibliotheca Pteridologia 1: 1-158.
  7. DIXIT, R.D. 1984. A Census of the Indian Pteridophytes. Botanical Survey of India, Howrah.
  8. DUTHIE, J.F. 1886. Kumaon. in Garden Chronicle. 25: 456-458.
  9. DUTHIE, J.F. 1906. Catalogue of the Plants of Kumaon and of the Adjacent Portions of Garhwal and Tibet, based on the collections made by Strachey and Winterbottom during the years 1846 to 1849 (Catalogue originally prepared in 1852 by Sir Richard Strachey). Lovell Reeve & Co., Limited, London.
  10. FRASER-JENKINS, C.R. 2008. Endemics and Pseudo-Endemics in Relation to the Distribution Patterns of Indian Pteridophytes. Taiwania. 53(3): 264-292.
  11. FRASER-JENKINS, C.R. 2011. Nepal's little known pteridophytes, their distribution, geography and the hidden work of David Don. website version updated 1 Dec. 2010, on
  12. FRASER-JENKINS, C.R. AND C.S. DULAWAT 2009. A summary of Indian Cheilanthoid ferns and the discovery of Negripteris (Pteridaceae), an Afro-Arabian fern genus new to India. Fern Gaz. 18: 216-229.
  13. GIEGER, J.M.O., T.A. RANKER, J.M.R. NEALE AND S.T. KLIMAS 2007. Molecular biogeography and origins of the Hawaiian fern flora. Brittonia 59: 142-158.
  14. HOOKER, W.J. 1858. Species Filicum vol. 2. Pamplin, London.
  15. HOOKER, W.J. AND J.G. BAKER 1874. Synopsis Filicum, ed. 2. R. Hardwicke, London.
  16. HOPE, C.W. 1899-1904. The Ferns of North-West India. J. Bombay Nat. His. Soc. 12: 315-325, 527-538, 621-633 (1899); 13: 25-36, 236-251 (1900); 13: 443-461, 657-671 (1901), 14: 118-127, 252-266, 458-480 (1902), 14: 720-749,15: 78-111 (1903); 15: 415-419 (1904). Reprinted in bookform in Delhi with altered pagination.
  17. KATO, M. 1993. Biogeography of ferns: dispersal and vicariance. Journal of Biogeography 20: 265-274.
  18. KESSLER, M. 2010. Biogeography of Ferns. In Fern Ecology (eds.) Mehltreter, K. Walker, L. R. and Sharpe , J. M. pp. 22-60. Cambridge University Press, New York.
  19. KHOLIA, B.S. AND K. BHAKUNI 2009. Western Himalaya-a new range of distribution for a critically endangered fern Dryopsis manipurensis (Bedd.) Holttum & P.J. Edwards Bot. Surv. India. 51; 245-248.
  20. KHOLIA, B.S. AND N. PUNETHA 2002. Diplazium subsinuatum (Wall.ex Hook, et Grev) Tag. In Central Himalaya. Phytotatxonomy 2: 163-165.
  21. KHOLIA, B.S. AND N. PUNETHA 2003. The genus Colysis Presl. in western Himalaya. Phytotaxonomny 3: 119-123.
  22. KHOLIA, B.S. AND N. PUNETHA 2005. Notes on Distribution and Conservation of the rare Selaginella pulvinata (Hook.& Grev.) Maxim, in India. Indian Journal of Fore stary 28(4): 389-391.
  23. KHULLAR, S.P 1994. An illustrated fern flora of west Himalaya Vol I. International Book Distributors Dehradun.
  24. KHULLAR, S.P, Y.P.S. PANGTEY, S.S. SAMANT, R.S. RAWAL AND G.S. RAW AT 1991. Ferns of Nainital. Bishen Singh Mahendra pal Singh Dehradun.
  25. MORAN, R.C. AND A.R. SMITH 1993. Phytogeographic relationships between neotropical and African-Madagascan Pteridophytes. Brittonia. 53: 304-351.
  26. PANDE, H.C. AND PC. PANDE 2002. Pteridology in the Western Himalaya (Kumaon). Bishan Singh Mahendra Pal Singh, Dehradun.
  27. PANDE, H.C AND PC. PANDE 2003. An Illustrated Fern Flora oF Kumaon Himalaya 1. Bishan Singh Mahendra Pal Singh, Dehradun.
  28. PANGTEY, Y.P.S., C.R., FRASER-JENKINS, LALIT TIWARI AND KANCHAN UPTERTI 2010. Ferns of Nainital, W. Himalaya: Some changes and corrections. Indian Fern J. 152-177.
  29. PUNETHA, N, AND B.S. KHOLIA 1998. Pteridophytic flora of Kumaon: Distribution, Evolution and status. Indian Fern 7.15: 75-79.
  30. PUNETHA, N, AND B.S. KHOLIA 1988. Vittaria himalayensis Ching, A new record for western Himalaya. Curr. Sci 57: 346-348.
  31. PUNETHA, N, AND B.S. KHOLIA 1989. Addition to the Pteridophytic flora of Pithoragarh district of Kumaon (W. Himalaya). New Botanist. 36: 115-126.
  32. RANKER, T.A., S.K. FLOYD AND P.G. TRAPP 1994. Multiple colonization of Asplenium adiantum-nigrum on to the Hawai'ian archipelago. Evolution. 48: 1364-1370.
  33. Roux, J.P. 2001. Conspectus of southern African Pteridophyta. Southern African Botanical Diversity Network Report No. 13. SABONET, Pretoria. Published by The Project Coordinator, Southern African Botanical Diversity Network, c/o National Botanical Institute Private Bag X101, Pretoria, 0001 South Africa.
  34. SHING, K.H. AND S.K. WU 1990. Sinopteridaceae. In R.C. Ching and K.H. Shing (Ed.). Flora Republicae Popularis Sinicae 3(1). Scineice Press. Beijing. 97-173.
  35. SMITH, A.R. 1993. Phytogeographic principles and their use in understanding fern relationships. Journal of Biogeography 20: 255-264.
  36. SMITH, A.R. 2010. Appendix B-Index to genera of ferns and lycophytes in alphabetical order. In Fern Ecology (eds.) Mehltreter, K. Walker, L.R. and Sharpe, J.M. pp. 386-405. Cambridge University Press, New York.
  37. TRYON, R.M., A.F., TRYON, K.U. KRAMER, 1990. Pteridaceae. In: Kramer, K.U., Green, P.S. (Eds.), The Families and Genera of Vascular Plants. Vol. 1. Pteridophytes and Gymnosperms. Springer-Verlag, Berlin, pp. 230-256.
  38. VERMA, S.C. AND S.P KHULLAR, 1980. Ferns of Nainital (W. Himalaya) An updaed list. Fern Gaz. 12: 83-92.
  39. WAGNER, W. H. 1995. Evolution of Hawai'ian ferns and fern-allies in relation to their conservations status. Pacific Science. 49: 31-41.